gradle-play-publisher: Fail to upload signed APK

Describe the bug

We currently experiencing an issue with uploading signed apk to the Google Play. We use v1 signing. This is our configuration:

play {
  serviceAccountCredentials = file("${rootProject.projectDir}/service.json")
  // specifying dir will just upload things and avoid to build apk locally
  // mapping file should be in the same folder
  artifactDir = file("${apkFolder}")
  track = 'alpha'
}

This error response that we see on our CI:

com.google.api.client.googleapis.json.GoogleJsonResponseException: 403 Forbidden
  {
    "code" : 403,
    "errors" : [ {
      "domain" : "androidpublisher",
      "message" : "APK signature is invalid or does not exist. Error from apksigner: ERROR: JAR_SIG_NO_MANIFEST: Missing META-INF/MANIFEST.MF",
      "reason" : "apkNotificationMessageKeyInvalidSignature"
    } ],
    "message" : "APK signature is invalid or does not exist. Error from apksigner: ERROR: JAR_SIG_NO_MANIFEST: Missing META-INF/MANIFEST.MF"
  }

The strange thing - we can still upload file manually.

How To Reproduce

Run our CI pipeline.

Versions

• Gradle Play Publisher: 2.2.1
• Gradle Wrapper: 5.4.1
• Android Gradle Plugin: 3.4.1

Tasks executed

app:publishProductionApk

Expected behavior

We can upload apk with gradle as we can do it manually.